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Community Classification of the Pawnee National Grasslands, Shortgrass Steppe, USA Scott B. Franklin 1 1, Michael Schiebout 2, & Amber Brandt 1 School of Biological Sciences, University of Northern Colorado, Greeley, CO USA; 2 Department of Biology, Union University, Jackson, TN USA But from these immense prairies may arise one great advantage to the United States, viz., the restriction of our population to some certain limits, and thereby a continuation of the union. Our citizens being so prone to rambling, and extending themselves on the frontiers, will, through necessity, be constrained to limit their extent on the west to the borders of the Missouri and the Mississippi, while they leave the prairies, incapable of cultivation, to the wandering and uncivilized Aborigines of the country. Zebulon Pike Purpose Results Classification of vegetation provides an inventory to assess change, a common language to compare communities among regions, and a baseline for land stewardship decisions. Several initial efforts toward mapping and vegetation data collection are available for the Pawnee National Grasslands, but not a structured community analysis and subsequent mapping for the entire area. The goal of this study is to provide such a local community analysis and relate community types to environmental conditions. Subsequently, local types will be related to The National Vegetation Classification (USNVC) developed as a standard for the United States. NMS ordination revealed clear significant patterns of community types, and subsequent discriminant analyses differentiated community types mainly by Bouteloua-Yucca percent cover and site type (an index Bouteloua-Opuntia Pascopyrum smithii of the moisture gradient). Bouteloua Ridgetop The Pawnee National Grasslands is located in northeastern Colorado with the Shortgrass Steppe Ecoregion, and part of the Colorado Piedmont and High Plains physiographic regions. The fragmented area is gently rolling with few outcrops; 78,128 ha. Physiography 2.0 NMS Axis 2 Study Area 2.5 Mixed Carex-Eleocharis Sporobolus-Glycyrrhiza Bouteloua gracilis Bouteloua dactyloides decreasing cover 1.5 The mixed community included everything from dry riparian to rock outcrops harboring mixed prairie species, Pinus flexilis, and shrubs. 1.0 0.5 0.0 increasing moisture -0.5 Site 1 -1.0 -2 -1 0 1 2 NMS Axis 1 Euclidean Distance Clustering using Ward’s Method Site 5 Sedimentary cycles Climate Mean Max Daily Temp in Summer = 31°C Bouteloua-Yucca Small Rock Outcrop Community Bouteloua-Opuntia Slope Community Site 7 Mean PPT = 305-380 mm Pascopyrum smithii Community Bouteloua Open [low cover] Community Slightly increasing moisture from west to east Remnant Buffalo Wallow Cottonwood riparian Mixed Community Geology includes shale, sandstone, sand dunes, and gravel; soils include badlands, mollisols, entisols, and alfisols. Site 2 Carex-Eleocharis Wet Community Sprobolis airoides-Glycyrrhiza lepidota Community Methods Bouteloua gracilis Community The initial phase of this project used GIS map layers to develop an ecological land type classification that was used to stratify field plots, including geology, soils, elevation, and landscape location (Kupfer & Franklin 2000; see Figures below). Site 97 Bouteloua dactyloides Community Site 4 Site 11 Site 44 Site 33 Discriminant Analysis accuracy table; values are percent of plots correctly classified Plot sampling utilized the Carolina Vegetation Sampling method modular plots (Peet et al. 1998); ten 10X10 modules = 20X50 m plots. A total of 102 plots were collected. Cover of taxa was recorded in four of the ten modules using a modified Daubenmire cover scale. Trees were measured in all ten modules. Community Type Bouteloua-Yucca Bouteloua-Opuntia Pascopyrum smithii Bouteloua Ridgetop Mixed Carex-Eleocharis Sprobolus- Glycyrrhiza Bouteloua gracilis Bouteloua dactyloides Various multivariate techniques were used to classify the local community types and relate those types to abiotic parameters. t t 0.1 1.1 0.5 0.5 t 0.1 0.3 0.1 0.1 Community Types Bouteloua- Bouteloua Boutelousa Pascopyrum SporobolisOpuntia gracilis dactyloides smithii Mixed Glycrrhiza 10.9 41.7 12.9 13.2 3.2 2.7 1.3 1.8 2.2 0.2 3.6 15.6 9.2 40.5 7.2 This content downloaded on Sun, 24 Feb 2013 16:38:25 PM 0.7 All use subject to JSTOR Terms and Conditions 1.3 1.9 1.5 0.3 0.2 0.2 0.2 t t t 4.7 8.8 6.1 7.6 0.5 t 5.2 2.6 2.6 37.5 0.9 8.2 1.5 0.3 0.2 0.2 0.5 t 1.8 t t 0.5 t 1.5 2 1.3 1.4 t 0.5 t 3.3 1.7 0.3 t t 39.2 0.2 12.5 t t 1.9 2.5 0.4 5.8 0.1 5.8 t t 0.2 0.1 0.2 0.3 2.9 t 1.5 0.8 0.2 0.5 t t t 41.7 Pascopyrum smithii 12.5 8.33 100 5.9 8.3 4.8 94.1 16.7 16.7 14.3 8.3 4.8 33.3 SporobolusGlycyrrhiza 25.0 25.0 100 8.3 Bouteloua gracilis Bouteloua dactyloides 12.5 8.3 16.7 100 8.3 23.8 50.0 14.3 8.3 38.1 The local Community types provided above are just that, local, and represent variations on concepts of wider distribution. However, it is important to place our local community types into the national classification concepts to fully describe the types. Most of the communities are part of the Great Plains Shortgrass Prairie Group. A total of 299 species (~430 known) were witnessed in the 102 plots scattered throughout the Pawnee National Grasslands. Eights distinct (one mixed) community types were distinguished through clustering analysis. Species Bouteloua gracilis Yucca glauca Bouteloua dactyloides Aristida purpurea Elymus elymoides Rhus trilobata Opuntia polyacantha Pascopyrum smithii Artemesia frigida Artemesia filifolia Atriplex canescens Schizochirium scoparium Toxicodendron rydbergii Sporobolus airoides Glycyrrhiza lepidota Distichlis spicata Elymus canadensis Juncus balticus Taraxacum officinale Poa spp Carex spp Juncus spp Eleocharis spp Circium arvense BoutelouaOpuntia Fitting the Local Community Types to the USNVC Results Bouteloua BoutelouaOpen Yucca 2.3 15 0.7 6.8 1.1 4.8 0.2 4.5 0.1 2.5 0.7 1.9 0.2 1.6 2.2 1.1 0.3 0.7 t 0.3 0.1 0.3 1.1 0.3 0.1 0.1 0.2 BoutelouaYucca 62.5 8.33 Community Type Bouteloua Mixed CarexOpen Eleocharis 12.5 8.3 t t Community Types BoutelouaYucca CarexEleocharis 0.1 t 0.1 1.7 0.1 0.9 1.1 3.1 26 13.5 7.4 1.7 BoutelouaOpuntia Bouteloua Boutelousa Pascopyrum SporobolisCarexUSNVC Bouteloua Open gracilis dactyloides smithii Glycyrrhiza Eleocharis Bouteloua Bouteloua Bouteloua gracilisArtemesia gracilis-Buchloe gracilis-Buchloe Great Plains Shortgrass Prairie & Shrubland Buchloe dactyloides Gutierrezia frigidadactyloides dactyloides MG Xeric Soil sarothrae – Boutleloua Herbaceous Herbaceous Great Plains Shortgrass Prairie Group Herbaceous Yucca glauca gracilis Dwarf- Vegetation Vegetation Vegetation Dwarf-shrub shrub ? ? Pascopyrum Great Plains Mixedgrass Prairie & Shrubland smithiiMG Bouteloua Central Great Plains Mixed Grass Prairie gracilis Group Artemesia filifoliaBouteloua Great Plains Sand Grassland & Shrubland gracilis http://usnvc.org/explore-classification/ MG Shrubland Take-home Messages A vegetation gradient was found from open rock communities to wet prairies. Local communities were related to environmental factors, but also typical disturbances (i.e., prairie dogs, grazing). The relation of these local types to the USNVC is clear for most of the area and common vegetation, but the USNVC is lacking mesic prairie association descriptions.