Chapter 27 Opener Figure 21.1 The Evolution of Plants Figure 21.1 The Evolution of Plants (Part 2)
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Chapter 27 Opener Figure 21.1 The Evolution of Plants Figure 21.1 The Evolution of Plants (Part 2) Figure 21.1 The Evolution of Plants (Part 1) Figure 21.4 Alternation of Generations in Land Plants Figure 21.4 Alternation of Generations in Land Plants In-Text Art, Ch. 27, p. 557 Figure 27.1 Perfect and Imperfect Flowers Figure 27.1 Perfect and Imperfect Flowers (Part 1) Figure 27.1 Perfect and Imperfect Flowers (Part 2) Figure 27.1 Perfect and Imperfect Flowers (Part 3) Concept 27.1 Most Angiosperms Reproduce Sexually Angiosperm gametophytes are microscopic. Female (megagametophyte), or embryo sac arises from a megaspore. Consists of 7 cells: • 1 egg cell • 2 synergids (attract pollen tube and receive sperm) • 3 antipodal cells degenerate • 1 central cell with 2 polar nuclei Concept 27.1 Most Angiosperms Reproduce Sexually Male (microgametophytes), or pollen grains, arise from microspores. Consist of 2 cells: Generative cell divides by mitosis to form two sperm cells that participate in fertilization. Tube cell forms pollen tube that delivers the sperm to embryo sac. Figure 27.2 Sexual Reproduction in Angiosperms Figure 27.3 Self-incompatibility Figure 27.4 Double Fertilization Figure 27.4 Double Fertilization Figure 27.5 Angiosperm Fruits Figure 27.5 Angiosperm Fruits (Part 1) Figure 27.5 Angiosperm Fruits (Part 2) Figure 27.5 Angiosperm Fruits (Part 3) In-Text Art, Ch. 27, p. 561 (1) In-Text Art, Ch. 27, p. 561 (2) Concept 27.1 Most Angiosperms Reproduce Sexually With a partner, discuss the differences between sexual reproduction in angiosperms and animals. Consider, for example, the products of mitosis and meiosis and the alternation of haploid and diploid generations. Concept 27.1 Most Angiosperms Reproduce Sexually Draw a diagram of a flower and label all of the following: Anther Filament Ovule Sepal Stigma Carpel Ovary Petal Stamen Style [Note: If a real flower is available, see if you can identify all of these parts on the real flower, too.] Compare your drawing with your neighbors and discuss: 1. Did you draw a perfect or an imperfect flower? 2. Where is the male gametophyte? 3. Where is the female gametophyte? 4. Where is the sporophyte? 5. Which part will become the seed? 6. Which will become the rest of the fruit? Concept 27.1 Most Angiosperms Reproduce Sexually Which of the following correctly describes an imperfect flower? a. Has male and female sex organs b. Has both stamens and carpels c. Has only male or only female sex organs d. I don’t know. Concept 27.1 Most Angiosperms Reproduce Sexually Without looking at your notes, answer the following questions. 1. Is self-fertilization possible in dioecious plants? Why or why not? 2. How is self-fertilization prevented in monoecious plants? When you are finished, discuss your answers with a partner. Concept 27.1 Most Angiosperms Reproduce Sexually Draw a fruit and label the following: • The seed with the embryo • The ovary wall Then answer the following questions: 1. What part of the flower are you eating when you bite into an apple? 2. What are the two main functions of fruit? Figure 27.6 The Transition to Flowering Figure 27.6 The Transition to Flowering Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State Genes that determine the transition to floral meristems have been studied in Arabidopsis. Meristem identity genes LEAFY and APETALA1 initiate a cascade of gene expression. Floral organ identity genes: homeotic genes; products are transcription factors that determine whether cells in the floral meristem will be sepals, petals, stamens, or carpels. Figure 27.6 The Transition to Flowering (Part 1) Figure 27.6 The Transition to Flowering (Part 2) Figure 27.6 The Transition to Flowering (Part 3) Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State External cues that initiate gene expression for flowering: 1. Photoperiod (day length)—Some species flower only when days reach a specific length. • Short-day plants (SDPs) flower only when the day is shorter than a critical maximum. • Long-day plants (LDPs) flower only when the day is longer than a critical minimum. Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State Plants sense night length by measuring the ratio of Pfr to Pr. Day—more red light than far-red; by end of day most phytochrome is Pfr. At night Pfr is gradually converted back to Pr. The longer the night, the more Pr there is at dawn. A SDP flowers when ratio of Pfr to Pr is low at the end of the night; a LDP flowers when this ratio is high. Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State Phytochrome is located in the leaf. The signal for flowering must be a diffusible chemical that travels from the leaf to the shoot apical meristem. The diffusible chemical is the protein florigen. Figure 39.19 Structure of a phytochrome Figure 39.20 Phytochrome: a molecular switching mechanism Figure 27.7 Photoperiod and Flowering Figure 27.7 Photoperiod and Flowering Figure 27.8 Night Length and Flowering Figure 27.8 Night Length and Flowering Hormones and signaling determine the transition from the vegetative to the flowering state The experiments reported on in fig. 27.8 were repeated using a long-day plant (LDP) that normally flowers with a 16-hour photoperiod. The plant was subjected to the light regimes listed in the table. 1. List the light regimes that should have resulted in flowering. 2. Predict the effect of adding a brief period of far-red light in the middle of the dark period of each light regime. 3. Predict the effect of adding a brief period of red light in the middle of the dark period of each light regime. Apply the Concept, Ch. 27, p. 564 Figure 39.22 Photoperiodic control of flowering Figure 27.9 The Flowering Signal Moves from Leaf to Bud Figure 27.9 The Flowering Signal Moves from Leaf to Bud (Part 1) Figure 27.9 The Flowering Signal Moves from Leaf to Bud (Part 2) Figure 27.10 Molecular Biology of Flowering Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State The genes involved in flowering: FT (FLOWERING LOCUS T) codes for florigen. CO (CONSTANS) codes for a transcription factor that activates synthesis of FT; expressed in phloem companion cells. FD (FLOWERING LOCUS D) codes for a transcription factor that binds to FT in the shoot apical meristem. The complex activates promoters for meristem identity genes, such as APETALA1. Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State External cues that initiate gene expression for flowering: 2. Temperature Some plants flower after a period of cold temperatures (vernalization). Cold temperatures inhibit synthesis of FLC protein, a transcription factor that inhibits expression of FT and FD. Figure 27.10 Molecular Biology of Flowering Figure 27.10 Molecular Biology of Flowering (Part 1) Figure 27.10 Molecular Biology of Flowering (Part 2) Figure 35.36 The ABC hypothesis for the functioning of organ identity genes in flower development Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State With a partner, review the different types of meristems. Then answer the following questions: 1. What do shoot apical meristems produce? 2. What do shoot apical meristems become? 3. What do inflorescence meristems produce? 4. What does a floral meristem become? Concept 27.2 Hormones and Signaling Determine the Transition from the Vegetative to the Reproductive State What is the critical factor that induces flowering? a. Day length b. Night length c. Age of plant d. Red light exposure Concept 27.3 Angiosperms Can Reproduce Asexually Why is cloning a good reproductive method for some plant species? That is, why might asexual reproduction be adaptive? Concept 27.3 Angiosperms Can Reproduce Asexually What is it called when one of these two events occurs? 1. The megasporocyte in the ovule that is supposed to undergo meiosis fails to do so, resulting in a diploid egg cell, which then goes on to form an embryo and seed. 2. Diploid cells from the integument surrounding the embryo sac form a diploid embryo sac, which goes on to form an embryo and seed. Does seed and fruit development proceed? What does the plant produce? Concept 27.3 Angiosperms Can Reproduce Asexually Which term does not belong with the others? a. Vegetative reproduction b. Cloning c. Apomixis d. Sexual reproduction e. Asexual reproduction Concept 27.3 Angiosperms Can Reproduce Asexually Describe the importance and function of the vascular cambium in the grafting process. Figure 27.11 Vernalization In-Text Art, Ch. 27, p. 568 Figure 27.12 Vegetative Reproduction Figure 27.12 Vegetative Reproduction Figure 27.12 Vegetative Reproduction (Part 1) Figure 27.12 Vegetative Reproduction (Part 2) Figure 27.12 Vegetative Reproduction (Part 3) Figure 27.13 The Advantage of Asexual Reproduction by Apomixis Figure 27.13 The Advantage of Asexual Reproduction by Apomixis Figure 27.14 Grafting Figure 27.14 Grafting Figure 27.15 A Wild Relative of Poinsettia