Modelling combined effects of elevation, aspect and slope on species-presence and growth Albert R.
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Modelling combined effects of elevation, aspect and slope on species-presence and growth Albert R. Stage and Christian Salas Old ideas • French scientists modelled wine cork lengths on different sides of oak trees 50 years ago with: a·Cos(aspect) + b·Sin(aspect) • Beers, Dress and Wensel 40 years ago (1966) recommended a·Cos(aspect + phase shift) where phase shift for the adverse aspect was assumed to be = SW • Stage 30 years ago (1976) added an interaction with slope to represent white pine site index: slope·[a·Cos(aspect) + b·Sin(aspect)+ c] and thereby allowing the data to determine the phase shift. Trig Tricks • Stage(1976) is a generalization of Beers, Dress and Wensel (1966) because: y = b0 + b1s + b2·s·cos(α) + b3·s·sin(α) is identical to: y = b0 + b1·s + b b cos(α - β) for β = +arctan(b3/b2) if b2 > 0 or −arctan(b2/b3) if b3 >0. 2 2 2 3 Now what about Elevation? • Roise and Betters (1981) argued that optimum phase shift reverses between elevation extremes-- but omitted aspect/slope relations in their formulation. • Here we combine these concepts in terms of main effects of elevation with two elevation functions interacting with slope/aspect triplets. Introducing the two elevation/aspect interactions: • Behavior: – Sensitivity to elevation increases toward the extremes (contra Roise and Betters 1981) – Scale invariant – Linear model preferred Introducing the two elevation/aspect interactions: F1(elev)·slope·[a1·Cos(aspect) + b1Sin(aspect)+ c1] + F2(elev)·slope·[a2·Cos(aspect) + b2·Sin(aspect)+ c2] + d1·F3(elev) Some alternative pairs of functions: F1 (low) F2 (high) Constant = 1 Square of elevation elevation Square of elevation Log(elevation) Square of elevation Log (k·elevation)= Log (elev) + log(k) Square of elevation Challenging hypothesis with DATA! • Where there is agreement--- Classifying forest/non-forest in Utah Slope = 20% 0.15 Elev. =1750 m. Discriminant Non-forest >|< Forest 0.1 Elev. = 4000 m. 0.05 0 -0.05 -0.1 -0.15 -0.2 0 90 180 Aspect 270 360 Utah productivity MAI - Utah Slope =25% 60 Cu. Ft. 50 40 north south level 30 20 10 0 1.5 2 2.5 3 3.5 Elevation (m/1000) 4 5 6 Challenging hypothesis with DATA! • Where there is agreement--• And where there is not ! Douglas-fir Height Growth F1 = ln(elev), K F2 = elev2 55 853 m. 1219 m. Asymptote (m) 50 1768 m. 45 40 35 30 N E S Aspect W N Douglas-fir Height Growth 3 elevation classes 55 640-1082 m. 1083-1311 m. Asymptote (m) 50 1312- 2073 m. 45 40 Not an artifact ! 35 30 N E S Aspect W N So · · · ? • Proposed formulation consistent with ecological hypotheses concerning elevation-aspect-slope relations · · · • But allows data to define some surprises !